Cursos que dicta


  • Métodos filogenéticos y macroevolutivos



Grados Académicos

Ph.D., Organismic and Evolutionary Biology. Harvard University, Cambridge, Massachusetts, Estados Unidos de América.

M.A., Evolution, Ecology, and Population Biology. Washington University in St. Louis, St. Louis, Missouri, Estados Unidos de América.

M.A., Biology, Boston University, Boston, Massachusetts, Estados Unidos de América.


Líneas de investigación:

  • Biología evolutiva.
  • Métodos computacionales macroevolutivos y filogenéticos.
  • El desarrollo de paquetes R por el análisis de datos biológicos.
  • Ecología evolutiva de reptiles.


Producción científica


2020-2023 FONDECYT Regular 2020: ShinyPhytools: A web platform for phylogenetic comparative methods, Award Number 1201869.

2018-2021 ABI Development: Computational infrastructure for phylogenetic & macroevolutionary analysis in the R environment. U.S. National Science Foundation, Award Number 1759940. Investigador principal.

2017-2018 Dissertation Research: Adaptive responses to urbanization in the tropical lizard Anolis cristatellus. U.S. National Science Foundation, Award Number 1701706. Investigador principal, con co-investigadora K. Winchell.

2014-2020 CAREER: Phylogenetic tools for studying phenotypic diversification in the tree of life. U.S. National Science Foundation, Award Number 1350474.

2014-2017 Collaborative Research: Urban adaptation and its role in the success of biological invasion in Anolis U.S. National Science Foundation, Award Number 1354044. Investigador principal, en proyecto colaborativo con investigadores principales J. Kolbe, Univ. Rhode Island, y D. Warner, Auburn Univ.

2006-2007 Dissertation Research: The evolution and importance of genetic constraint in Anolis cristatellus. U.S. National Science Foundation, Award Number 0608291. Co-investigador, con investigador principal J. Losos.


Publicaciones (*: WOS)

Winchell, K. M., K. P. Schliep, D. L. Mahler, and L. J. Revell (2020) Phylogenetic signal and evolutionary correlates of urban tolerance in a widespread neotropical lizard clade. Evolution 74: 1274-1288.*

Reynolds, R. G., J. J. Kolbe, R. E. Glor, M. López-Darias, C. V. Gómez Pourroy, A. S. Harrison, K. de Queiroz, L. J. Revell, and J. B. Losos (2020) Phylogeographic and phenotypic outcomes of brown anole colonization across the Caribbean provide insight into the beginning stages of an adaptive radiation. Journal of Evolutionary Biology 33:468-494.*

Quach, Q. N., R. G. Reynolds, and L. J. Revell (2020) Historical allopatry and secondary contact or primary intergradation in the Puerto Rican Crested Anole, Anolis cristatellus, on Vieques Island in the Caribbean. Biological Journal of the Linnean Society 129: 114-127.*

Prado-Irwin, S. R., L. J. Revell, and K. M. Winchell (2019) Variation in tail morphology across urban and forest populations of the crested anole (Anolis cristatellus). Biological Journal of the Linnean Society 128:632-644.*

Revell, L. J (2019) learnPopGen: an R package for population genetic simulation and numerical analysis. Ecology and Evolution 9: 7896-7902.*

Winchell, K. M., D. Briggs, and L. J. Revell (2019) Perils of city life: Patterns of injury and fluctuating asymmetry in urban lizards.
Biological Journal of the Linnean Society 126: 276-288.*

Bawa, K. S., T. Ingty, L. J. Revell, and K. N. Shivaprakash (2019) Correlated evolution of flower size and seed number in flowering plants (monocotyledons). Annals of Botany 123: 181-190.*

Revell, L. J., K. Schliep, E. Valderrama, and J. E. Richardson ( 2018) Graphs in phylogenetic comparative analysis: Anscombe’s quartet revisited. Methods in Ecology and Evolution 9: 2145-2154. *

Winchell, K. M., I. Maayan, J. R. Fredette, and L. J. Revell (2018) Linking locomotor performance to morphological shifts in urban lizards. Proceedings of the Royal Society, B 285: 20180229. *

Revell, L. J ( 2018) Comparing the rates of speciation and extinction between phylogenetic trees. Ecology and Evolution 8: 5303-5312. *

Revell, L. J., L. E. González-Valenzuela, A. Alfonso, L. A. Castellanos-García, C. E. Guarnizo, and A. J. Crawford (2018) Comparing evolutionary rates between trees, clades, and traits. Methods in Ecology and Evolution 9: 994-1005. *

Winchell, K. M., E. J. Carlen, A. R. Puente-Rolón, and L. J. Revell (2018) Divergent habitat use of two urban lizard species. Ecology and Evolution 8: 25-35. *

Reynolds, R. G., T. R. Strickland, J. J. Kolbe, B. G. Falk, G. Perry, L. J. Revell, and J. B. Losos (2017) Archipelagic genetics in a widespread Caribbean anole. Journal of Biogeography 44: 2631-2647. *

Arcila, D., R. Vari, J. W. Armbruster, M. L. J. Stiassny, K. D. Kol, M. H. Sabaj, J. Lundberg, L. J. Revell, G. Ortí, and R. Betancur-R (2017) Genome-wide gene genealogy interrogation advances resolution of recalcitrant groups in the Tree of Life. Nature Ecology & Evolution 1: 0020. *

Tyler, R. K., K. M. Winchell, and L. J. Revell (2016) Tails of the city: Caudal autotomy in the tropical lizard, Anolis cristatellus, in urban and natural areas of Puerto Rico. Journal of Herpetology 50: 435-441. *

Reynolds, R. G., D. C. Collar, S. A. Pasachnik, M. L. Niemiller, A. R. Puente-Rolón, and L. J. Revell (2016) Ecological specialization and morphological diversification in Greater Antillean boas. Evolution 70: 1882–1895. *

Winchell, K. M., R. G. Reynolds, S. Prado-Irwin, A. R. Puente-Rolón, and L. J. Revell (2016) Phenotypic shifts in urban areas in the tropical lizard, Anolis cristatellus. Evolution 70: 1009-1022. *

Harrison, A. S., L. J. Revell, and J. B. Losos (2015) Correlated evolution of microhabitat, morphology, and behavior in West Indian Anolis lizards: A test of the habitat matrix model. Behaviour 152: 1187-1207. *

Revell, L. J., D. L. Mahler, R. G. Reynolds, and G. J. Slater (2015) Placing cryptic, recently extinct, or hypothesized taxa into an ultrametric phylogeny using continuous character data: A case study with the lizard Anolis roosevelti. Evolution 69: 1027-1035. *

Reynolds, R. G., A. R. Puente-Rolón, R. Platenberg, K. Tyler, P. J. Tolson, and L. J. Revell (2015) Large divergence and low diversity suggest genetically informed conservation strategies for the endangered Virgin Islands Boa (Chilabothrus monensis). Global Ecology and Conservation 3: 487-502. *

Collar, D. C., P. C. Wainwright, M. E. Alfaro, L. J. Revell, and R. S. Mehta (2014) Biting disrupts integration to spur skull evolution in eels. Nature Communications 5: 5505. *

Stuart, Y. E., T. S. Campbell, R. G. Reynolds, P. A. Hohenlohe, L. J. Revell, and J. B. Losos (2014) Rapid evolution of a native species following invasion by a congener. Science 346: 463-466. *

Revell, L. J., and S. A. Chamberlain (2014) Rphylip: An R interface for PHYLIP. Methods in Ecology and Evolution 5: 976-981. *

Reynolds, R. G., A. R. Puente-Rolón, M. Barandiaran, and L. J. Revell (2014) Hispaniolan boa (Chilabothrus striatus) on Vieques Island, Puerto Rico. Herpetology Notes 7: 121-122.

Revell, L. J (2014) Ancestral character estimation under the threshold model from quantitative genetics. Evolution 68: 743-759. *

Reynolds, R. G., M. L. Niemiller, and L. J. Revell (2014) Toward a tree-of-life for the boas and pythons: Multilocus species-level phylogeny with unprecedented taxon sampling. Molecular Phylogenetics and Evolution 71: 201-213. *

Revell, L. J. (2013) Two new graphical methods for mapping trait evolution on phylogenies. Methods in Ecology and Evolution 4: 754-759. *

Mahler, D. L., T. Ingram, L. J. Revell, and J. B. Losos (2013) Exceptional convergence on the macroevolutionary landscape in island lizard radiations. Science 341: 292-295. *

Reynolds, R. G., M. L. Niemiller, S. B. Hedges, A. Dornburg, A. R. Puente-Rolón and L. J. Revell (2013) Molecular phylogeny and historical biogeography of West Indian boid snakes (Chilabothrus). Molecular Phylogenetics and Evolution 68: 461-470. *

Puente-Rolón , A. R., R. G. Reynolds, and L. J. Revell. Preliminary genetic analysis supports cave populations as targets for conservation in the endemic endangered Puerto Rican boa (Boidae: Epicrates inornatus). PLoS ONE 8: e63899. *

Muñoz, M. M., N. G. Crawford, T. J. McGreevy, N. J. Messana, R. D. Tarvin, L. J. Revell, R. M. Zandvliet, J. M. Hopwood, E. Mock, A. L. Schneider, and C. J. Schneider (2013) Divergence in coloration and the evolution of reproductive isolation in the Anolis marmoratus species complex. Molecular Ecology 22: 2668-2682. *

Reynolds, R. G., A. R. Puente-Rolón, R. N. Reed, and L. J. Revell (2013) Genetic analysis of a novel invasion of Puerto Rico by an exotic constricting snake. Biological Invasions 15: 953-959. *

Revell, L. J. (2013) A comment on the use of stochastic character maps to estimate evolutionary rate variation in a continuously valued trait. Systematic Biology 62: 339-345. *

Revell, L. J. and R. G. Reynolds. (2012) A new Bayesian method for fitting evolutionary models to comparative data with intraspecific variation. Evolution 66: 2697-2707. *

Revell, L. J (2012) phytools: An R package for phylogenetic comparative biology (and other things). Methods in Ecology and Evolution 3: 217-223. *

Slater, G. J., L. J. Harmon, D. Wegmann, P. Joyce, L. J. Revell, and M. E. Alfaro (2012) Fitting models of continuous trait evolution to incompletely sampled comparative data using Approximate Bayesian Computation. Evolution 66: 752-762. *

Sanger, T. J., L. J. Revell, J. J. Gibson-Brown, and J. B. Losos (2012) Repeated modification of early limb morphogenesis programs underlies the evolution of relative long bone length variation among Anolis lizards. Proceedings of the Royal Society, B 279: 739-748. *

Revell, L. J., D. L. Mahler, P. R. Peres-Neto, and B. D. Redelings (2012) A new phylogenetic method for identifying exceptional phenotypic diversification. Evolution 66: 135-146. *

Kolbe, J. J., L. J. Revell, B. Szekely, E. D. Brodie III, and J. B. Losos (2011) Convergent evolution of phenotypic integration and its alignment with morphological diversification in Caribbean Anolis Evolution 65: 3608-3624. *

Revell, L. J. (2010) Phylogenetic signal and linear regression on species data. Methods in Ecology and Evolution 1: 319-329. *

Mahler, D. L., L. J. Revell, R. E. Glor, and J. B. Losos (2010) Ecological opportunity and the rate of morphological evolution in the diversification of Greater Antillean anoles. Evolution 64: 2731-2745. *

Lindenfors, P., L. J. Revell, and C. L. Nunn (2010) Sexual dimorphism in primate aerobic capacity: A phylogenetic test. Journal of Evolutionary Biology 23: 1183-1194. *

Johnson, M. A., L. J. Revell, and J. B. Losos (2010) Behavioral convergence and adaptive radiation: Effects of habitat use on territorial behavior. Evolution 64: 1151-1159. *

Lovely, K. R., D. L. Mahler, and L. J. Revell (2010) The rate and pattern of tail autotomy in five species of Puerto Rican anoles. Evolutionary Ecology Research 12: 67-88. *

Revell, L. J., D. L. Mahler, J. R. Sweeney, M. Sobotka, V. E. Fancher, and J. B. Losos (2010) Nonlinear selection and the evolution of variances and covariances for continuous characters in an anole. Journal of Evolutionary Biology 23: 407-421. *

Revell, L. J. (2009) Size-correction and principal components for interspecific comparative studies. Evolution 63: 3258-3268. *

Revell, L. J. and D. C. Collar (2009) Phylogenetic analysis of the evolutionary correlation using likelihood. Evolution 63: 1090-1100. *

Revell, L. J., L. J. Harmon, and D. C. Collar (2008) Phylogenetic signal, evolutionary process, and rate. Systematic Biology 57: 591-601. *

Revell, L. J. (2008) On the analysis of evolutionary change along single branches in a phylogeny. American Naturalist 172: 140-147. *

Revell, L. J., and L. J. Harmon (2008) Testing quantitative genetic hypotheses about the evolutionary rate matrix for continuous characters. Evolutionary Ecology Research 10: 311-331. *

Revell, L. J., and A. S. Harrison (2008) PCCA: A program for phylogenetic canonical correlation analysis. Bioinformatics 24: 1018-1020. *

Johnson, M. A., M. Leal, L. Rodríguez Schettino, A. Chamizo Lara, L. J. Revell, and J. B. Losos (2008) A phylogenetic perspective on foraging mode evolution in West Indian Anolis Animal Behaviour 75: 555-563. *

Revell, L. J., M. A. Johnson, J. A. Schulte II, J. J. Kolbe, and J. B. Losos (2007) A phylogenetic test for adaptive convergence in rock-dwelling lizards. Evolution 61: 2898-2912. *

Revell, L. J. (2007) Testing the genetic constraint hypothesis in a phylogenetic context: A simulation study. Evolution 61: 2720-2727. *

Revell, L. J. (2007) The G matrix under fluctuating correlational mutation and selection. Evolution 61: 1857-1872. *

Revell, L. J., L. J. Harmon, R. B. Langerhans, and J. J. Kolbe. (2007) A phylogenetic approach to determining the importance of constraint on phenotypic evolution in the neotropical lizard Anolis cristatellus. Evolutionary Ecology Research 9: 261-282. *

Revell, L. J., L. J. Harmon, and R. E. Glor (2005) Underparameterized model of sequence evolution leads to bias in the estimation of diversification rates from molecular phylogenies. Systematic Biology 54: 973-983. *


Capítulos de libros

Revell, L. J (2014) Graphical methods for visualizing comparative data on phylogenies. Chapter 4 in Modern phylogenetic comparative methods and their application in evolutionary biology: Concepts and practice (L. Z. Garamszegi ed.). ISBN 978-3-662-43550-2.

Software científico

Revell, L. J. 2011-20. phytools: Phylogenetic tools for comparative
biology (and other things). R package version 0.7-70.

Revell, L. J. and S. A. Chamberlain. 2013-20. Rphylip: An R interface
for PHYLIP. R package version 0.1-29.

Revell, L. J. and C. L. Revell. 2017-20. ishihara: Ishihara color-vision
test. R package version 0.3.

Revell, L. J. 2012-19. learnPopGen: Population genetic simulations &
numerical analysis. R package version 1.0.5.

Revell, L. J. 2017. physketch: Drawing phylogenetic objects in R
environment. R package version 0.1.

Más información

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